REYNOLDS A.J., JAHODA C.A., Hair matrix epidermal cells confer follicle-inducing capabilities on dermal sheath & high passage papilla cells. // Development., vol.122(10)., p.3085-3094., 1996

2. In vitro, trichocytes, isolated from plucked human scalp HFs were propagated on feeder layers of post-mitotic human DF. When grown as 'surface epithelia' in vitro [as 2-D culture, on dermal equivalents exposed to air], trichocytes organized into stratified epithelia largely reminiscent of E with regard to both tissue architecture and localization of epidermal differentiation. Keratinization was virtually complete when trichocytes grown on collagen gels with DF were transplanted onto nude mice; this was apparent by tissue organization, expression of K1 and K10 and the nearly regular epidermal localization of involucrin. In addition, the deposition of basement membrane components [laminin, type IV collagen, bullous pemphigoid antigen] at the epithelium-collagen interface further increased and was more regular in transplants than in vitro.
Trichocytes embedded in Matrigel together with DF developed large spheroidal structures with inward-directed differentiation and all the epidermal markers found in 'surface' cultures, while only small keratinizing spheroids formed without HDF.

LIMAT A., BREITKREUTZ D., THIEKOETTER G., NOSER F., HUNZIKER T., BRATHEN L.R., FUSENIG N.E. Phenotypic modulation of human hair matrix cells (trichocytes0 by environmental influence in vitro & in vivo. // Epithelial Cell biol., vol.2(2)., p.55-65., 1993
LIMAT A., BREITKREUTZ D., STARK H-J., HUNZINKER T., THIKOETTER G., NOSER F., FUSENIG N.E., Experimental modulation of the differentiated phenotype of keratinocytes from epidermis & hair follicle outer root sheath 7 matrix cells. // 1991 Annals of the New York academy of sciences . vol 642,p.125-147